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Ecosystems often contain large numbers of species and those species are themselves highly variable; but how is this diversity maintained? Our group focuses on this very large question by using a variety of experimental systems and a combination of observation, modelling and experiment. Currently, important themes are plant growth modelling, plant-soil feedbacks, and the impact of herbivores and hemiparasites. Regular collaborators include Drew Purves (Microsoft Research), Mark Rees (University of Sheffield) and Daniel Kliebenstein (University of California, Davis).